The translocation of organic solutes in sieve tube members is supported by
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P-Protein bodies and callose form blocking plugs. These bodies remain along the walls of sieve tube elements. P-Protein bodies are assigned some in the translocation of food material in the sieve tubes but is not universally accepted.
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According to mass flow hypothesis the transport of organic solutes takes place from source to sink, this transport also depends on metabolic energy. According to cytoplasmic strearming hypothesis (which was given by de Vries, 1885) the transport of organic solutes takes place by combination of diffusion and cytoplasmic streaming. Cytoplasmic streaming carry organic solutes from one and to the other end of sieve tube. P-proteins has a role as defence against phloem feeding insects and sealing of damaged sieve tubes.
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