Who is reflecting his father's gnarled disease??
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Syntactically speaking, his lesson is parenthetical, or nonrestrictive, information. Analyzing the sentence, it would be possible to drop it: a father's gnarled disease, his lesson, from his desk. So, while it may be understood that he is reciting his lesson, what the sentence is saying at its core is that he is reciting the gnarled disease.
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