write a note on bond fision or cleavage
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EMBRYOLOGY LECTURE NOTES -2
CLEAVAGE AND TYPES
The process of cleavage reamains one of the earliest mechanical activity in the conversion of a
single celled egg into a multicellular embryo. It is initiated by the sperm during fertilization.
However in parthenogenetic eggs cleavage can commence without the influence of fertilization.
The process of cleavage or cellulation happens through repeated mitotic divisions. These divisions
result in cells called blastomeres. In later stages of development the blastomeres occupy different
regions and differentiate into several types of body cells.
The first cleavage of frog’s egg was observed by Swammerdam in 1738. The entire process of
cleavage in frog’s egg was studied by Prevost and Dumas in 1824. With the development of
microscopes cleavages and further stages were observed in the eggs of sea urchin, star fishes,
amphioxus and hen’s eggs.
From all these studies it has become clear that all divisions in cleavage are mitotic. The mitotic
process is very rapid. In the eggs of sea urchin division of the blastomeres can be observed every 30
minutes.
As the cleavage progresses the resultant daughter cells, namely the blastomeres get reduced in size.
During cleavage there is no growth in the blastomeres. The total size and volume of the embryo
remains the same.
The cleavages result in a compact mass of blastomeres called morula. It gets transformed into
blastula. While the wall of the blastula is called the blastoderm, the central cavity is called the
blastocoel.
The planes of cleavage
An egg can be divided from different planes during cleavage. Depending on the position of the
cleavage furrow the planes of cleavage are named.
1. Meridional plane: The plane of cleavage lies on the animal vegetal axis. It bisects both the poles
of the egg. Thus the egg is divided into two equal halves.
Meridional cleavage