Question

affinities of echinoderms with chordates​

Answer 1

The origin and affinities of chordates and echinoderms have long been a subject of interesting speculation. Relationships between creatures are postulated mainly on grounds of comparative anatomy and development. For existing creatures, the methods of comparative physiology and serology are also of great importance. In any comparison also involving fossil form, it is important to remember that any evolutionary change does not affect the whole existing stock. Such a change could occur in one locality only and the existing stock might still survive in other areas. Thus it is that we have such ”living fossils’’ as the kingcrab, the coelacanth, cycas and Equisetum. A primitive existing form may be very closely related to the ancestor of more advanced existing forms.

From the available evidence, it seems fairly certain that chordates and echinoderms are related. In the first place, we have the striking resemblance between the tornaria larva of Balanoglossus and the auricularia larva of echinoderms. Both are strikingly different from the characteristic trochophore larva of annelid and molluscs. Also, during the development of lower chordates and echinoderms, the mesoderm is formed in the same manner, by budding of pouches dorso-laterally from the archenteron. The cavities of these pouches constitute the beginning of the coelom. This is different from the manner in which mesoderm and coelom. This is in annelids, arthropods and molluscs. Both in echinoderms and chordates, thecentral nervous system develops from a dorsal strip of epiblast. A final point of comparison lies in the type of phosphagen present in the muscle.

In chordates the phosphate. However, in Balanoglossus and in a few echinoderms, both types of phosphagen are found. The chordate and echinoderm stocks must have diverged very early in animal evolution; fossils of both types are found in early Palaeozoic rocks. It is often suggested that we must look for chordate origins in the coelenterates. There seems to be no evidence that Rhabdopleura, a deep-sea hemichordate, shows features in its colony formation which show a strong resemblance to the same process in the fossil graptolites. Unfortunately, Rhabdopleura has very slender qualifications for inclusion in the chordate. It has no visceral clefts or dorsal hollow nerve cord, and its notochord is a structure like that of Balanoglossus. Here again we may possibly consider that both the graptolits and the ancestral hemichordate were offshoots which diverged early from the coelenterate stock. Comparison of Amphioxus has been made with two Silurian fossils of Jamoytius Kerwoodi. These fossils show a soft- bodies animal with a distinct notochord and with the same general arrangement of intestine and myotomes as aree found in Amphioxus. But Jamoytius had undoubted eyes. It seems that by the Silurian period, the graptolites, Jamoytius Ostracoderms related to the present lampreys, and the bony fishes had already been evolved.

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Answer 2

Affinities Of Echinoderms With Chordates

The origin and affinities of Echinoderms With Chordates have extremely existed as the subject of intriguing assumptions. Relationships between creatures are postulated primarily for purposes of comparable anatomy and growth. According to the accessible information, it appears relatively certain that echinoderms have some relation with chordates.

• Firstly, we have an extraordinary resemblance between the tornaria larva of Balanoglossus and the auricularia larva of echinoderms. Both are extraordinarily distinct from the ordinary trochophore larva of annelids and molluscs.

• Secondly, during the growth of lower chordates and echinoderms, the mesoderm was established in a similar way, by growing pouches dorsolaterally from the archenteron. The basins of these pouches comprise the outset of the coelom. This is unusual from how mesoderm and coelom are in annelids, arthropods and molluscs.

• Thirdly, both in echinoderms and chordates, the central nervous system evolves from a dorsal piece of the epiblast.

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