Question

what factors may be responsible for huge losses during storage?

Answer 1

the factor responsible for storage loss are biotic and abiotic
biotic = fungi ,bacteria ,pests, insects
abiotic = moisture and temperature of storage place

Answer 2

Both biotic and abiotic factors are responsible for the loss of stored grains. Abiotic factors include improper temperature of store house, humidity ofair and high moisture content of the grains. Improper containers used for storage also cause loss of grains.